New Taxa
and major taxonomic revisions by HERMANN W. PFEFFERKORN
1 new family, 6 new genera, 16 new species, 9 new combinations, 2 major revisions,
2 species named after HWP
Status: December 2021
Taxonomy and nomenclature are not pursued for their own sake but whenever it becomes necessary to clear up taxonomic and/or nomenclatural problems. This work is the basis of reliable data sets for the solutions of major questions, i.e. correct interpretations of global change, paleoclimate, paleoecology, and global correlations.
NEW FAMILY
Nystroemiaceae Wang et Pfefferkorn
Wang, J. & Pfefferkorn, H.W. 2009. Nystroemiaceae, a new family of Permian gymnosperms from China with an unusual combination of features.
Proceedings of the Royal Society B, v. 277, p. 301-309.
A broad-leaved gymnosperm family of Permian age that combines very advanced with very ancient features. These trees had a modern growth form (similar to Ginkgo) but bicornute ovules that were born in bifurcate and pinnate structures reminiscent of early Carboniferous pteridosperms.
NEW GENERA
Highlight indicates name of person honored
Circulipuncturites Labandeira, Wang, Zhang, Bek et Pfefferkorn
Wang, J., Labandeira, C.C., Zhang, G., Bek, J., & Pfefferkorn, H.W. 2009. Permian Circulipuncturites discinisporis Labandeira, Wang, Zhang, Bek et Pfefferkorn gen. et spec. nov. from China, an ichnotaxon of a punch-and-sucking insect on noeggerathialean spores. Review of Paleobotany and Palynology, v. 156, p. 277-282.
Otzinachsonia Cressler et Pfefferkorn
Cressler, W.L. & Pfefferkorn, H.W. 2005. A Late Devonian isoetalean lycopsid, Otzinachsonia beerboweri gen. et sp. nov., from north-central Pennsylvania, U.S.A. American Journal of Botany, v. 92, p. 1131-1140.
Oclloa Erwin, Pfefferkorn et Alleman
Obandotheca Erwin, Pfefferkorn et Alleman
Erwin, D.M., Pfefferkorn, H.W. & Alleman, V. 1994. Early seed plants in the Southern Hemisphere: I. Associated ovulate and microsporangiate organs from the Carboniferous of Peru. Review of Paleobotany and Palynology, v. 80, p. 19-38.
We honored the director of the Paracas Natural Preserve at the time of publication.
Artisophyton Pfefferkorn
Pfefferkorn, H.W. 1976. Pennsylvania tree fern compressions Caulopteris, Megaphyton, and Artisophyton gen, nov. in Illinois. Illinois State Geological Survey Circular 492, 31 pp.
Honored is Edmund Tyrell Artis (1789-1847) who did an early, careful study of Carboniferous tree fern stems.
Kankakeea Pfefferkorn
Pfefferkorn, H.W. 1973. Kankakeea gen. nov., buds for vegetative reproduction in Carboniferous ferns. Palaeontologische Zeitschrift, v. 47, p. 143-151.
NEW SPECIES
Nemejcopteris haiwangii Pšenička, Wang, Bek, Pfefferkorn, Opluštil, Weiming, Votočková Frojdová et Libertín
Pšenička, J., Wang, J., Bek, J., Pfefferkorn, H.W., Opluštil, S., Weiming Z., Votočková Frojdová, J., Libertín, M., 2021. A zygopterid fern with fertile and vegetative parts in anatomical and compression preservation from the earliest Permian of Inner Mongolia, China. Review of Palaeobotany and Palynology, v. 294, https://doi.org/10.1016/
We honor Haiwang Zhang whose interest in paleobotanical samples and help with paleobotanical research in Inner Mongolia, China, was of great importance for the success of the project.
Palaeostachya guanglongi Liu, Pšenička, Bek, Wan, Pfefferkorn et Wang
Liu, L., Pšenička, J., Bek, J., Wan, M., Pfefferkorn, H.W., Wang, J., 2020. A whole calamitacean plant Palaeostachya guanglongii from the Asselian (Permian) Taiyuan Formation in the Wuda Coalfield, Inner Mongolia, China. Review of Palaeobotany and Palynology, v. 294, https://doi.org/10.1016/
We honor Guanglong Shen for his contributions to Late Paleozoic paleobotany of China.
Aphlebia hvistendahliae Wang et Pfefferkorn
Wang, J., Wan, M., Pfefferkorn, H.W. 2014. Aphlebia hvistendahliae sp. nov. from the early Permian Wuda Tuff Flora, Inner Mongolia. Review of Palaeobotany and Palynology, v. 210, p. 69-76.
We honored Mara Hvistendahl, a science writer, who showed interest in our work, visited the field area, and found the specimen of the new species.
Circulipuncturites discinisporis Labandeira, Wang, Zhang, Bek et Pfefferkorn
Wang, J., Labandeira, C.C., Zhang, G., Bek, J., & Pfefferkorn, H.W. 2009. Permian Circulipuncturites discinisporis Labandeira, Wang, Zhang, Bek et Pfefferkorn gen. et spec. nov. from China, an ichnotaxon of a punch-and-sucking insect on noeggerathialean spores. Review of Paleobotany and Palynology, v. 156, p. 277-282.
Paratingia wudensis Wang, Pfefferkorn et Bek 2009
Wang, J., Pfefferkorn, H.W., & Bek, J. Paratingia wudensis sp. nov., a whole noeggerathialean plant preserved in an air fall tuff of earliest Permian age (Inner Mongolia, China). American Journal of Botany, v. 96, p. 1676-1689.
Marattia aganzhenensis Yang, Wang et Pfefferkorn
Yang, Shu, Wang, Jun, & Pfefferkorn, H.W. 2008 Marattia aganzhenensis sp. nov. from the Lower Jurassic Daxigou Formation of Lanzhou, Gansu, China. International Journal of Plant Science, v. 169, p. 473-482.
Otzinachsonia beerboweri Cressler et Pfefferkorn
Cressler, W.L. & Pfefferkorn, H.W. 2005. A Late Devonian isoetalean lycopsid, Otzinachsonia beerboweri, gen. et sp. nov., from north-central Pennsylvania, U.S.A. American Journal of Botany, v. 92, p. 1131-1140.
We honored James Richard (Dick) Beerbower (1927-2010) for his investigations into the development of early terrestrial ecosystems.
Discinites baculiformis Wang, Pfefferkorn, Feng et Shen
Yuania wudensis Wang, Pfefferkorn, Feng et Shen
Wang J., Pfefferkorn, H.W., Feng, Zhou, & Shen Guanglong. 2004. A new species of Discinites (Noeggerathiales) associated with a new species of Yuania from the Lower Permian of Inner Mongolia, China. International Journal of Plant Science, v. 165, p. 1107-1119.
Neuralethopteris sergiorum Goubet, Pfefferkorn et Gillespie
Goubet, P., Pfefferkorn, H.W. & W.H. Gillespie. 2000. Neuralethopterids (Trigonocarpalean Pteridosperms) from the Early Pennsylvanian of eastern North America. PaleoBios, v. 20(3), p. 11-37.
We honored three great paleobotanists by using the Latin second case plural: Sergio Archangelsky (Argentina), Sergius H. Mamay (1921-2008, USA), and Sergei V. Meyen (1935-1987, Russia).
Oclloa cesariana Erwin, Pfefferkorn et Alleman
Obandotheca laminensis Erwin, Pfefferkorn et Alleman
Erwin, D.M., Pfefferkorn, H.W. & Alleman, V. 1994. Early seed plants in the Southern Hemisphere: I. Associated ovulate and microsporangiate organs from the Carboniferous of Peru. Review of Paleobotany and Palynology, v. 80, p. 19-38.
We honored Silvia N. Césari the eminent Argentinian paleobotanist.
Lesleya cheimarosa Leary et Pfefferkorn
Rhodeopteridium phillipsii Leary et Pfefferkorn
Leary, R.L. & H.W. Pfefferkorn. 1977. An early Pennsylvanian flora with Megalopteris and Noeggerathiales from west-central Illinois. Illinois State Geological Survey Circular 500, 77 pp.
We honored Tom L. Phillips (1931-2018) in whose lab we were working when we began the study of the Spencer Farm Flora.
Kankakeea grundyi Pfefferkorn
Pfefferkorn, H.W. 1973. Kankakeea gen. nov., buds for vegetative reproduction in Carboniferous ferns. Palaeontologische Zeitschrift, v. 47, p. 143-151.
Myriotheca arnoldii Pfefferkorn ex Pfefferkorn, Peppers et Phillips
Pfefferkorn, H.W., Peppers, R.A. & T.L. Phillips. 1971. Some fern-like fructifications and their spores from the Mazon Creek compression flora of Illinois (Pennsylvanian). Illinois State Geological Survey Circular 463, 29 pp.
We honored Chester A. Arnold (1901-1977) who among other areas did pioneering work on late Paleozoic compression floras.
NEW COMBINATIONS
Tomiodendron peruvianum (Gothan) Pfefferkorn et Alleman ex Iannuzzi et Pfefferkorn
Iannuzzi, R. & H.W. Pfefferkorn. 2002. A pre-glacial, warm-temperate floral belt in Gondwana (Late Visean, Early Carboniferous). PALAIOS, v. 17, p. 571-590.
Neuralethopteris pocahontas (D. White) Goubet, Pfefferkorn et Gillespie
Neuralethopteris smithsii (Lesquereux) Goubet, Pfefferkorn et Gillespie
Neuralethopteris pentias (D. White) Goubet, Pfefferkorn et Gillespie
Neuralethopteris biformis (Lesquereux) Goubet, Pfefferkorn et Gillespie
Goubet, P., Pfefferkorn, H.W. & W.H. Gillespie. 2000. Neuralethopterids (Trigonocarpalean Pteridosperms) from the Early Pennsylvanian of eastern North America. PaleoBios, v. 20(3), p. 11-37.
Artisophyton goldenbergii (Weiss) Pfefferkorn
Artisophyton mclayi (Lesquereux) Pfefferkorn
Pfefferkorn, H.W. 1976. Pennsylvania tree fern compressions Caulopteris, Megaphyton, and Artisophyton gen, nov. in Illinois. Illinois State Geological Survey Circular 492, 31 pp.
Kankakeea thallyformis (Lesquereux) Pfefferkorn
1973 Pfefferkorn, H.W. Kankakeea gen. nov., buds for vegetative reproduction in Carboniferous ferns. Palaeontologische Zeitschrift, v. 47, p. 143-151.
Stellatheca ornata (Lesquereux) Pfefferkorn ex Pfefferkorn, Peppers et Phillips
1971 Pfefferkorn, H.W., Peppers, R.A. & T.L. Phillips. Some fern-like fructifications and their spores from the Mazon Creek compression flora of Illinois (Pennsylvanian). Illinois State Geological Survey Circular 463, 29 pp.
MAJOR REVISIONS
Genus Neuralethopteris in the United States
The genus Neuralethopteris belongs to the gymnosperms (pteridosperms or “seed ferns”). Species of the genus Neuralethopteris are common in late Mississippian and early Pennsylvanian strata and several of them serve as index species for specific time intervals. The family becomes suddenly extinct at the boundary between the Langsettian and Duckmantian stages (Westphalian A-B boundary) in the Euramerican Realm. A few authors had recognized that the genus occurred in North America but no systematic study and formal introduction existed before this publication.
Goubet, P., Pfefferkorn, H.W. & W.H. Gillespie. 2000. Neuralethopterids (Trigonocarpalean Pteridosperms) from the Early Pennsylvanian of eastern North America. PaleoBios, v. 20(3), p. 11-37.
Systematic position and morphometrics of the genus Discopteris (fern)
Pfefferkorn, H.W. 1978. Revision der Sphenopteriden Discopteris karwinensis
STUR, Discopteris vuellersi STUR und der Gattung Discopteris STUR.
Argumenta Palaeobotanica v. 5, p. 167-193.
REMOVAL OF INVALID TAXA
I have not counted how many species or genera I have declared nomina nuda or invalid for other reasons. In one particular Carboniferous genus over 60 species had been given a name in the literature but only 17 were valid species. There were synonyms, nomina nuda, and dubious species that had to be removed. Similar results would be found in other groups after a critical revision based on type specimens and a complete set of the literature of figured specimens. Also necessary is a critical attitude combined with modern measuring and statistical methods to determine what is objectively distinguishable.
Not counted was a publication that was superseded by the publication of Labandeira et al. 2009
NEW TAXA
named after
HERMANN W. PFEFFERKORN
Duodemidia pfefferkornii Cressler, Prestianni et LePage 2010
Cressler, W.L. III, Prestianni, C., LePage, B.A. 2010. Late Devonian spermatophyte diversity and paleoecology at Red Hill, north-central Pennsylvania, USA. International Journal of Coal Geology, v. 83, p. 91-102.
Sigillaria pfefferkornii D'Antonio, Boyce et Wang 2020
D'Antonio, M.P., Boyce, C.K., Wang, J. 2020. Two new species of Sigillaria Brongniart from the Wuda Tuff (asselian: Inner Mongolia, China) and their implications for lepidodendrid life history reconstruction. Review of Paleobotany and Palynology, https://doi.org/10.1016/j.revpalbo.2020.104203